Ari Benjamin

Cold Spring Harbor Laboratory.

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I study how and why neural networks learn what they do. I see neural networks both as models of learning in the brain and as intelligent systems in their own right, and I have spent the past decade training them and moving between these two views.

Much of my work is theory. I have shown that efficient neural codes emerge naturally from gradient descent (Nature Communications, 2022), studied how to measure and regularize networks in the space of functions they compute (ICLR, 2019), and recently showed that a network can be understood as a Bayesian ensemble of its tangent functions — a lens that reframes catastrophic forgetting and points toward new algorithms for continual learning (NeurIPS, 2024).

I also like to build. I designed and trained TissueFormer, a transformer that extends single-cell foundation models to predict tissue-level phenotypes from populations of single cells (BMC Bioinformatics, 2026), and have worked on uncertainty-aware objectives for post-training language models.

The brain is still my favorite learning machine. I am especially drawn to what neural networks don’t yet capture: neuromodulators like serotonin, the wild diversity of cell types, and how brains keep learning for a lifetime without overwriting what they know. Lately this has me thinking about belief formation and the self-consistency of knowledge in large language models — what is learned in context versus in weights, and what complementary learning systems might look like in modern AI. These strike me as questions that could be understood analytically, and I’d love to see that happen.

Along the way I have co-organized a COSYNE workshop on why networks learn what they do, co-authored lecture notes on connectionist theory with Andrew Saxe, Jay McClelland, and colleagues at the Gatsby Unit’s Analytical Connectionism summer school, and reviewed for NeurIPS, ICML, and Nature Communications.

I am currently a postdoctoral fellow in the laboratory of Tony Zador at Cold Spring Harbor Laboratory. I completed my PhD with Konrad Kording at the University of Pennsylvania.

selected publications

  1. NeurIPS
    Continual learning with the neural tangent ensemble
    Ari Benjamin, Christian-Gernot Pehle, and Kyle Daruwalla
    Advances in Neural Information Processing Systems, 2024
  2. Nat. Comm.
    Efficient neural codes naturally emerge through gradient descent learning
    Ari S Benjamin, Ling-Qi Zhang, Cheng Qiu, Alan A Stocker, and Konrad P Kording
    Nature Communications, 2022
  3. PLOS CB
    A role for cortical interneurons as adversarial discriminators
    Ari S Benjamin and Konrad P Kording
    PLOS Computational Biology, 2023
  4. PMLR
    An Introduction to Connectionist Theories of Semantic Cognition
    Ari S Benjamin, Anna-Lea Beyer, Marianne De Heer Kloots, Jaedong Hwang, Hajer Karoui, and 6 more authors
    In Analytical Connectionism School, 2026
  5. BMC
    Tissueformer: extending single-cell foundation models to predict population-level phenotypes
    Ari S Benjamin and Anthony M Zador
    BMC Bioinformatics, 2026